The thrust of a jet or propeller doesn't make a plane fly, the forward motion that thrust imparts pushes the wings through the air creating lift which leads to flight. Assuming it is possible for a treadmill to move fast enough to create enough friction in the tires, wheel bearings, etc. that the thrust couldn't overcome to create forward movement then the airplane would not take off. It would have to be extremely fast since that would take a hell of a lot of speed to generate that much friction, there couldn't be enough wind to create lift without forward motion, and of course it couldn't be a VTOL aircraft.
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